Ecology · Biodiversity · CNRS

Nicolas Mouquet

Senior research scientist at CNRS, based at MARBEC and the University of Montpellier. Research at the interface of metacommunities, biodiversity, ecosystem functioning, functional biogeography, rarity, aesthetics of biodiversity, and nature futures.

Research overview

Ecology across scales

I am primarily an ecologist, with complementary experience in microbiology and evolution. I worked on some of the earliest metacommunity models, demonstrating how dispersal among local communities may affect both local and regional diversity. These models are important for formalizing our understanding of metacommunity dynamics.

I have also incorporated elements of network ecology (trophic biogeography & metaweb), ecosystem functioning (metaecosystems) and ecophylogenetics into this framework. I developed integrated approaches to understand how coexistence mechanisms shape the relationship between biodiversity and ecosystem functioning. I have incorporated elements of evolutionary dynamics mainly by developing experimental approaches with microorganisms.

I am now interested in disentangling the factors that shape the different facets of biodiversity, functional rarity, functional biogeography and the relationship between humans and nature through the aesthetics of biodiversity and the Nature for Future Framework. I have directed or collaborated in several reviews and conceptual syntheses.

Research themes

Explore topic pages

Publications

List of publications

  1. [136]
    Citation self-awareness for a fairer academic publishing landscape.

    Beck B., Annasawmy P., Birre B., Busana B., Casajus C., Coux C., Marino M., Mouquet N., Nicvert L., Oliveira B.F., Petit-Cailleux C., Tortosa A., Unkule M., Vagnon C., Veytia D. (2026) Citation self-awareness for a fairer academic publishing landscape. BioScience, 2026, biag028. doi.org/10.1093/biosci/biag028

  2. [135]
    Marine protected areas marginally offset anthropogenic declines in tropical reef fish contributions to nature and people.

    Flandrin, U., Mouquet N., Loiseau, N., Hautecoeur, C., Maire, E., McLean, M.,Sanchez, L., Clausius, E., Stuart-Smith, R., Edgar, G., Albouy, C., Cinner, J. & Mouillot, D. (2026). Marine protected areas marginally offset anthropogenic declines in tropical reef fish contributions to nature and people. Nature Ecology & Evolution, doi.org/10.1038/s41559-026-03008-2

  3. [134]
    Spatial Insurance of Distinct Ecological Functions.

    Mouquet N., Mahaut L., Thuiller W., Auber A., Casajus C., Enquist B.J., Gauzere, P., Loiseau N., D. Mouillot D., Munoz F., Villeger and Violle C. (2025). Spatial Insurance of Distinct Ecological Functions. Ecology Letters, 2025; 28:e70266, doi.org/10.1111/ele.70266

  4. [133]
    The science of cuteness: where teddy bears beauty, comfort and care converge.

    Blanc N., Tribot A.S., Brassac T., and Mouquet N. (2025). The science of cuteness: where teddy bears beauty, comfort and care converge. The Journal of Positive Psychology, doi.org/10.1080/17439760.2025.2538890

  5. [132]
  6. [131]
    Benthic communities on restored coral reefs confer equivalent aesthetic value to healthy reefs

    Alisa, C.A.G., Razak, T.B., Mouquet N.*,Graham A.J, HemingsonC. R. ,Mouillot D., Damayanti, L. Prasetya M. E. , Maulana P. B., Hamka A. , Dwiyanto A., Abeng A. T., Madjid, Irwan, R. E., Hidayat A., Lakota H., Parrangan C. V., Pratama A. M. A., Hamzah, Suandar, Subhan B., Zamani N. P., Vida R. T. & Lamont, T. A. C. (2025). Benthic communities on restored coral reefs confer equivalent aesthetic value to healthy reefs Scientific Reports 15, 20790. doi.org/10.1038/s41598-025-06373-3

  7. [130]
    Conserving the beauty of the worlds reef fish assemblages

    McLean M.*, Mouillot D., Langlois J., Arif S., Bejarano S., Casajus C., Edgar G.J., Flandrin U., Guilhaumon F., Judah A.B., Loiseau N., MacNeil M.A., Maire E., Stuart-Smith R.D., & Mouquet N.* (2025). Conserving the beauty of the worlds reef fish assemblages Proc. Natl. Acad. Sci. U.S.A. 122 (25) e2415931122, doi.org/10.1073/pnas.2415931122 (* These authors contributed equally to this study)

  8. [129]
    Assessing the ecological and aesthetic effectiveness of restoration interventions on coralligenous reefs

    Casoli E., Raschetti S., Mouquet N., Ventura D., Mancini G., Penna M., Andrea Belluscio A. and Ardizzone, G. (2025). Assessing the ecological and aesthetic effectiveness of restoration interventions on coralligenous reefs Restoration Ecology, e70092, doi.org/10.1111/rec.70092

  9. [128]
    Functional diversity shapes the stability of reef fish biomass under global change

    Mahaut L., Loiseau L., Villeger S., Auber A., Hautecoeur C., Maire A., Mellin C., Mouquet N., Stuart-Smith R., Violle C., and Mouillot D. (2025). Functional diversity shapes the stability of reef fish biomass under global change Proceedings of the Royal Society B, 292, 20250252, doi/10.1098/rspb.2025.0252

  10. [127]
    Danger versus fear: A key to understanding biophobia

    Zeller K., Mouquet N., Garcia C., Dezecache G., Maille A., Duboscq J., Morino J.L., & Bonnet X. (2025). Danger versus fear: A key to understanding biophobia People and Nature, doi: 10.1002/pan3.70009

  11. [126]
    Fish communities can simultaneously contribute to nature and people across the worlds tropical reefs

    Flandrin U., Mouillot D., Albouy C., Bejarano S., Casajus C., Cinner J., Edgar G., Ghilardi M, Leprieur L., Loiseau L., MacNeil A., Maire E., McLean M., Parravicini V., Pellissier L., Schiettekatte N., Stuart-Smith R., Villeger S. & Mouquet N. (2024) Fish communities can simultaneously contribute to nature and people across the worlds tropical reefs One Earth 7, 1772-1785. DOI:10.1016/j.oneear.2024.09.011/

  12. [125]
    The socioeconomic and environmental niche of protected areas reveals global conservation gaps and opportunities

    Mouillot D., Velez L., Albouy C., Casajus N., Claudet J., Delbar V., Devillers R., Letessier T.B., Loiseau N., Manel M., Mannocci L., Meeuwig J., Mouquet N., Nuno A., OConnor L., Parravicini P., Renaud J., Seguin R.,Troussellier T. & Thuiller W. (2024) The socioeconomic and environmental niche of protected areas reveals global conservation gaps and opportunities Nature Communications 15:9007. DOI:10.1038/s41467-024-53241-1

  13. [124]
    Inferring the extinction risk of marine fish to inform global conservation priorities

    Loiseau N., Mouillot D., Velez L., Seguin R., Casajus N., Coux C., Albouy C., Claverie T., Duhamet A., Fleure V., Langlois J., Villeger S. & Mouquet N. (2024) Inferring the extinction risk of marine fish to inform global conservation priorities PLOS Biology 22, e3002773. DOI:10.1371/journal.pbio.3002773

  14. [123]
    Low human interest for the most at-risk reef fishes worldwide

    Mouquet N., Langlois J., Casajus N., Auber A., Flandrin U., Guilhaumon F., Loiseau N., McLean M., Aurore R., Stuart Smith R.D. & Mouillot D. (2024) Low human interest for the most at-risk reef fishes worldwide Science Advances, 10,eadj9510. DOI:10.1126/sciadv.adj9510. This paper was featured in Science Advances by Becca Franks: Vulnerable fishes, inattentive humans

  15. [122]
    Divergent responses of pelagic and benthic fish body-size structure to remoteness and protection from humans

    Letessier T.B., Mouillot D., Mannocci L., Jabour Christ H., Elamin Mohammed E., Mohamed Elamin S., Friedlander A.M., Alex Hearn. A., Juhel J.B., Ring Kleiven A., Moland E., Mouquet N., Nillos-Kleiven P.J., Sala E., Thompson C.D.H., Velez L., Vigliola, L., Meeuwig J.J. (2024) Divergent responses of pelagic and benthic fish body-size structure to remoteness and protection from humans Science, 383, 976:982. DOI:10.1126/science.adi7562

  16. [121]
  17. [120]
    Early signals of Posidonia oceanica meadows recovery in a context of wastewater treatment improvements

    Bockel T., Marre G., Delaruelle G., Agel N., Boissery P., Guilhaumon F., Mouquet N., Mouillot D. & Deter J. (2024) Early signals of Posidonia oceanica meadows recovery in a context of wastewater treatment improvements Marine Pollution Bulletin 201, DOI:10.1016/j.marpolbul.2024.116193

  18. [119]
    chessboard: An R package for creating network connections based on chess moves

    Casajus N., Rievrs Borges E., Delaruelle G., Tabacchi E., Fried G. & Mouquet N. (2023) chessboard: An R package for creating network connections based on chess moves Journal of Open Source Software, 8(90), 5753, DOI:10.21105/joss.05753

  19. [118]
    Anchoring pressure and the effectiveness of new management measures quantified using AIS data and a mobile application

    Bockel T., Marre G., Delaruelle G., Holon F., Boissery P., Blandin A., Mouquet N., Deter J. (2023) Anchoring pressure and the effectiveness of new management measures quantified using AIS data and a mobile application Marine Pollution Bulletin 195, DOI:10.1016/j.marpolbul.2023.115511

  20. [117]
    Trait-environment associations diverge between native and alien breeding bird assemblages on the world's oceanic islands

    Rault C,Leprieur F, Barbaro L, Kreft H, Mouquet N, Papaix J, Cagan HS, Violle C & Barnagaud JY (2023) Trait-environment associations diverge between native and alien breeding bird assemblages on the world's oceanic islands Global Ecology and Biogeography, DOI:10.1111/geb.13729

  21. [116]
    The ecological causes of functional distinctiveness in communities

    Munoz F, Klausmeier CA, Gauzere P, Kandlikar PG, Litchman E, Mouquet N, Ostling A, Thuiller W, Algar AC, Auber A, Cadotte MW, Delalandre L, Denelle P, Enquist BJ, Fortunel C, Grenie M, Loiseau N, Mahaut L, Maire A, Mouillot D, Pimiento C, Violle C, and Kraft NJB (2023) The ecological causes of functional distinctiveness in communities Ecology Letters, DOI:10.1111/ele.14265

  22. [115]
  23. [114]
    Artificial Intelligence and Social Networks Can Help to Protect the Famous Dugong

    Mannocci L, Carcaillet F, Chaumont M, Givord-Coupeau B, Iovan C, Mouquet N , Raffard E, Ravera J, Peyre Y, Ponce A, Vigliola L, Villon S and Mouillot D (2022) Artificial Intelligence and Social Networks Can Help to Protect the Famous Dugong Front. Young Minds. 10:828597. doi: 10.3389/frym.2022.828597

  24. [112]
    Mitigation Strategies to Improve Reproducibility of Poverty Estimations From Remote Sensing Images Using Deep Learning.

    Machicao J, A Ben Abbes, L Meneguzzi, PLP Correa, A Specht, R David, G Subsol, D Vellenich, R Devillers, S Stall, N Mouquet, M Chaumont, L Berti-Equille, and D Mouillot (2022) Mitigation Strategies to Improve Reproducibility of Poverty Estimations From Remote Sensing Images Using Deep Learning. Earth and Space Science 9:e2022EA002379.

  25. [111]
    The aesthetic value of reef fishes is globally mismatched to their conservation priorities.

    Langlois J*, Guilhaumon F, Baletaud F, Casajus N, De Almeida Braga C, Fleure V, Kulbicki K, Loiseau N, Mouillot D, Renoult JP, Stahl A, Stuart Smith RD, Tribot AS & N, Mouquet* (2022) The aesthetic value of reef fishes is globally mismatched to their conservation priorities. PLoS Biol 20(6): e3001640. doi:10.1371/journal.pbio.3001640 (* These authors contributed equally to this study)

  26. [110]
    Plant community impact on productivity: Trait diversity or key (stone) species effects?

    P Brun, C Violle, D Mouillot, N, Mouquet, B J Enquist, F Munoz, T Munkemuller, A Ostling, N E Zimmermann, W Thuiller (2022) Plant community impact on productivity: Trait diversity or key (stone) species effects? Ecology Letters doi:10.1111/ele.13968

  27. [109]
    Functionally distinct tree species support long-term productivity in extreme environments

    L Delalandre, P Gauzere, W Thuiller, M Cadotte, N, Mouquet, D Mouillot, F Munoz, P Denelle, N Loiseau, X Morin, C Violle (2022) Functionally distinct tree species support long-term productivity in extreme environments Proceedings of the Royal Society B doi:10.1098/rspb.2021.1694

  28. [108]
    Niche breadth affects bacterial transcription patterns along a salinity gradient.

    A Rain Franco, N, Mouquet C Gougat Barbera, T Bouvier, S Beier (2021) Niche breadth affects bacterial transcription patterns along a salinity gradient. Molecular ecology doi:10.1111/mec.16316

  29. [107]
    Maintaining momentum for collaborative working groups in a post-pandemic world.

    Srivastava SD, Winter M, Gross LJ, Metzger JP, Baron JS, Mouquet N, Meagher T, Halpern BS & Pillar VD (2021) Maintaining momentum for collaborative working groups in a post-pandemic world. Nature Ecology & Evolution doi:10.1038/s41559-021-01521-0

  30. [106]
    Leveraging social media and deep learning to detect rare megafauna in video surveys.

    Mannocci L, Villon S, Chaumont M, Guellati N, Mouquet N, Iovan C, Vigliola L & Mouillot D (2021) Leveraging social media and deep learning to detect rare megafauna in video surveys. Conservation Biology, 11, doi:10.1111/cobi.13798

  31. [105]
    An integrated approach to estimate aesthetic and ecological values of coralligenous reefs.

    Langlois J, Guilhaumon F, Bockel T, Boissery P, De Almeida Braga C, Deter J, Holon F, Marre G, Tribot AS & Mouquet N (2021) An integrated approach to estimate aesthetic and ecological values of coralligenous reefs. Ecological Indicators, 129, 107935, doi:10.1016/j.ecolind.2021.107935

  32. [104]
    The dimensionality and structure of species trait spaces.

    Mouillot D, Loiseau N, Grenie M, Algar AC, Allegra M, Cadotte MW, Casajus N, Denelle P, Gueguen N, Maire A, Maitner M, McGill BJ, McLean M, Mouquet N, Munoz F, Thuiller T, Villeger S, Violle C and, Auber A (2021) The dimensionality and structure of species trait spaces. Ecology Letters, doi:10.1111/ele.13778

  33. [103]
    Global distribution and conservation status of ecologically rare mammal and bird species.

    Loiseau N*, Mouquet N*, Casajus N, Grenie M, Gueguen M, Maitner B, Mouillot D, Ostling A, Renaud J, Tucker C, Velez L, Thuiller W, and Violle C. (2020) Global distribution and conservation status of ecologically rare mammal and bird species. Nature Communication, 11, 5071 doi:10.1038/s41467-020-18779-w. (* These authors contributed equally to this study)

  34. [102]
    How community adaptation affects biodiversity ecosystem functioning relationships.

    Aubree F, David P, Jarne P, Loreau M, Mouquet N and Calcagno V(2020) How community adaptation affects biodiversity ecosystem functioning relationships. Ecology Letters, 23, 1263-1275 doi:10.1111/ele.13530

  35. [101]
    Coral Reef Fish: Not Just a Matter of Beauty!

    Boudin E, Carcaillet F, Tribot A, Carabeux Q, Deter J, Claverie T, Villeger S and Mouquet N (2020) Coral Reef Fish: Not Just a Matter of Beauty! Front. Young Minds. 8:13. doi:10.3389/frym.2020.00013

  36. [100]
    Species diversity and composition drive the aesthetic value of coral reef fish assemblages.

    Tribot, A.S, Deter, J., Claverie, T., Guillhaumon, F., Villeger, S., & Mouquet, N. (2019). Species diversity and composition drive the aesthetic value of coral reef fish assemblages. Biology letters, 15, 20190703, doi:10.1098/rsbl.2019.0703

  37. [99]
    How consideration of islands has inspired mainstream ecology: links between the theory of island biogeography and some other key theories.

    Warren B.H., R.E. Ricklefs, C. Thebaud, D. Gravel, & Mouquet, N. (2019). How consideration of islands has inspired mainstream ecology: links between the theory of island biogeography and some other key theories. In: Goldstein, M.I. and DellaSala, D.A. Encyclopedia of the Worlds Biomes: Islands and Atolls. Elsevier, Oxford.

  38. [98]
    Intraspecific and interspecific trait variability in tadpole metacommunities from the Brazilian Atlantic rainforest.

    Xavier Jordani, M. Mouquet, N., Casatti, L., Menin, M., de Cerqueira Rossa Feres, D. & Helene Albert, C.H. (2019). Intraspecific and interspecific trait variability in tadpole metacommunities from the Brazilian Atlantic rainforest. Ecology and evolution 9 (7), 4025-4037, doi:10.1002/ece3.5031

  39. [97]
    On the relative importance of space and environment in farmland bird community assembly.

    Henckel, L., Meynard, C.N., Devictor, V., Mouquet, N.& Bretagnolle, V. (2019). On the relative importance of space and environment in farmland bird community assembly. PloS one 14 (3), e0213360, doi:10.1371/journal.pone.0213360

  40. [96]
    Integrating the aesthetic value of landscapes and biological diversity.

    Tribot, A.S, Deter, J, & Mouquet, N. (2018). Integrating the aesthetic value of landscapes and biological diversity. Proceedings of the Royal Society B, 285, 20180971, doi:0.1098/rspb.2018.0971

  41. [95]
    Confronting species aesthetics with ecological functions in coral reef fish.

    Tribot, A.S, Carabeux, Q., Deter, J., Claverie, T., Villeger, S., & Mouquet, N. (2018). Confronting species aesthetics with ecological functions in coral reef fish. Scientific Reports, 8, 11733, doi:10.1038/s41598-018-29637-7

  42. [94]
  43. [93]
    Size evolution in microorganisms masks trade-offs predicted by the growth rate hypothesis.

    Gounand, I., T. Daufresne, D. Gravel, C. Bouvier, T. Bouvier, M. Combe, C. Gougat-Barbera, F. Poly, C. Torres-Barcelo, Mouquet, N. (2018).Size evolution in microorganisms masks trade-offs predicted by the growth rate hypothesis. Proceedings of the Royal Society B-Biological Sciences 283, doi:10.1098/rspb.2016.2272

  44. [92]
    A Common Toolbox to Understand, Monitor or Manage Rarity? A Response to Carmona et al.

    Violle, C., Thuiller, W., Mouquet, N., Munoz F., Kraft N.J.B., Cadotte M.W., Livingstone S.W., Grenie M., Mouillot D. (2017). A Common Toolbox to Understand, Monitor or Manage Rarity? A Response to Carmona et al. Trends in Ecology and Evolution doi:10.1016/j.tree.2017.10.001

  45. [91]
    Diversity spurs diversification in ecological communities.

    Calcagno, V., Jarne, P., Loreau, M., Mouquet, N. and David, P. (2017). Diversity spurs diversification in ecological communities. Nature Communication, doi:10.1038/ncomms1581

  46. [90]
    Functional Rarity: The Ecology of Outliers.

    Violle, C., Thuiller, W., Mouquet, N., Munoz F., Kraft N.J.B., Cadotte M.W., Livingstone S.W., Grenie M., Mouillot D. (2017). Functional Rarity: The Ecology of Outliers. Trends in Ecology and Evolution,32,356-367, doi:10.1016/j.tree.2017.02.002

  47. [89]
    How life-history traits affect ecosystem properties: effects of dispersal in meta-ecosystems.

    Massol, F., Altermatt, F., Gounand, I., Gravel, D., Leibold, M.A. and Mouquet, N. (2017). How life-history traits affect ecosystem properties: effects of dispersal in meta-ecosystems. Oikos,126,532-546, doi:10.1111/oik.03893

  48. [88]
  49. [87]
  50. [86]
    Taxonomic and functional diversity increase the aesthetic value of coralligenous reefs.

    Tribot, A.S.,Mouquet, N., Villeger, S., Raymond, M., Hoff, F., Boissery, P., Holon, F., Deter, J. (2016). Taxonomic and functional diversity increase the aesthetic value of coralligenous reefs. Scientific Reports, doi:10.1038/srep34229

  51. [85]
  52. [84]
  53. [83]
    Predicting biotic interactions and their variability in a changing environment.

    Kadowaki, K., Barbera, C.G., Godsoe, W., Delsuc, F.,Mouquet, N.(2016). Predicting biotic interactions and their variability in a changing environment. Biology letters, doi:10.1098/rsbl.2015.1073

  54. [82]
    MANGAL - making ecological network analysis simple.

    Poisot, T., Baiser, B.,Dunne, J.A.,Kefi, S.,Massol, F.,Mouquet, N.,Romanuk, T.N.,Stouffer, D.B.,Wood, S.A.,Gravel, D.(2016). MANGAL - making ecological network analysis simple. Ecography, 39, 384-390, doi:10.1111/ecog.00976

  55. [81]
  56. [80]
    A theory for species co-occurrence in interaction networks.

    Cazelle, K., Araujo, M.B., Mouquet N., and Gravel, D. (2016). A theory for species co-occurrence in interaction networks. Theoretical Ecology, 9, 039-048, doi:10.1007/s12080-015-0281-9

  57. [79]
  58. [78]
    Predictive Ecology in a Changing World.

    Mouquet N. , Lagadeuc, Y., Devictor, V., Doyen, L., Duputie, A., Eveillard, D., Faure, D., Garnier, E., Gimenez, O., Huneman, P., Jabot, F., Jarne, P., Joly, D., Julliard, R., Kefi, S., Kergoat, G.J., Lavorel, S., Le Gall, L., Meslin, L., Morand, S., Morin, X., Morlon, H., Pinay, G., Pradel, R., Schurr, F.M., Thuiller, W. and Loreau, M. (2015). Predictive Ecology in a Changing World. Journal of Applied Ecology, 52, 1293-1310, doi:10.1111/1365-2664.12482

  59. [77]
  60. [76]
    Representing taxonomic, phylogenetic and functional diversity: new challenges for Mediterranean marine-protected areas.

    Guilhaumon, F., Albouy, C., Claudet, J., Velez, L., Lasram, F.B., Tomasini, J.A., Douzery, E.J.P., Meynard, C.N., Mouquet N., Troussellier, M., Araujo, M.B., Mouillot, D. (2015). Representing taxonomic, phylogenetic and functional diversity: new challenges for Mediterranean marine-protected areas. Diversity and Distributions, 21, 175-187, doi:10.1111/ddi.12280

  61. [75]
  62. [74]
    Metacommunity speciation models and their implications for diversification theory.

    Hubert, N., Calcagno, V., Etienne, R.S. and Mouquet N. (2015). Metacommunity speciation models and their implications for diversification theory. Ecology Letters, 18, 864-881, doi:10.1111/ele.12458

  63. [73]
    Islands as model systems in ecology and evolution: prospects fifty years after MacArthur-Wilson.

    Warren, B.H., Simberloff, D., Ricklefs, R.E., Aguilee, R., Condamine, F.L., Gravel, D., Morlon, H., Mouquet N., Rosindell, J., Casquet, J., Conti, E., Cornuault, J., Fernandez-Palacios, J.M., Hengl, T., Norder, S.J., Rijsdijk, K.F., Sanmartin, I., Strasberg, D., Triantis, K.A., Valente, L.M., Whittaker, R.J., Gillespie, R.G., Emerson, B.C., Thebaud, C. (2015). Islands as model systems in ecology and evolution: prospects fifty years after MacArthur-Wilson. Ecology Letters, 18, 200-217, doi:10.1111/ele.12398

  64. [72]
    Empirical Evaluation of Neutral Interactions in Host-Parasite Networks.

    Canard, E.F., Mouquet N., Mouillot, D., Stanko, M., Miklisova, D. & Gravel, D. (2014). Empirical Evaluation of Neutral Interactions in Host-Parasite Networks. The American Naturalist, 183, 468-479, doi:10.1086/675363

  65. [71]
    The Paradox of Enrichment in Metaecosystems.

    Gounand, I., Mouquet N., Canard, E., Guichard, F., Hauzy, C. and Gravel, D. (2014). The Paradox of Enrichment in Metaecosystems. The American Naturalist, 184, 752-763, doi:10.1086/678406

  66. [70]
    Multifaceted diversity-area relationships reveal global hotspots of mammalian species, trait and lineage diversity.

    Mazel, F., Guilhaumon, F., Mouquet N., Devictor, V., Gravel, D., Renaud, J., Cianciaruso, M.V., Loyola, R., Diniz-Filho, J.A.F., Mouillot, D., and Thuiller, W. (2014). Multifaceted diversity-area relationships reveal global hotspots of mammalian species, trait and lineage diversity. Global Ecology and Biogeography, 23, 836-847, doi:10.1111/geb.12158

  67. [69]
    Estimates of species extinctions from species–area relationships strongly depend on ecological context.

    Matias M.G., Gravel D., Guilhaumon F., Desjardins-Proulx P., Loreau M., Munkemuller T. and Mouquet N. (2014). Estimates of species extinctions from species–area relationships strongly depend on ecological context. Ecography 37, 1-12, doi:10.1111/j.1600-0587.2013.00448.x

  68. [68]
    Asynchrony of taxonomic, functional and phylogenetic diversity in birds.

    Monnet, A.C., Jiguet, F., Meynard, C.N., Mouillot, D., Mouquet N., Thuiller, W. and Devictor, V. (2014). Asynchrony of taxonomic, functional and phylogenetic diversity in birds. Global Ecology and Biogeography, 23, 780-788, doi:10.1111/geb.12179

  69. [67]
    Spatial mismatch of phylogenetic diversity across three vertebrate groups and protected areas in Europe.

    Zupan, L., Cabeza, M., Maiorano, L., Roquet, C., Devictor, V., Lavergne, S., Mouillot, D., Mouquet N., Julien Renaud, J., and Thuiller, W. (2014). Spatial mismatch of phylogenetic diversity across three vertebrate groups and protected areas in Europe. Diversity and Distributions, 20, 674-685, doi:10.1111/ddi.12186

  70. [66]
    Confronting the Paradox of Enrichment to the Metacommunity Perspective.

    Hauzy C., Nadin G., Canard E., Gounand I. Mouquet N. and Ebenman B. (2013). Confronting the Paradox of Enrichment to the Metacommunity Perspective. Plos One, 8, e82969, doi:10.1371/journal.pone.0082969

  71. [65]
    Disentangling the role of connectivity, environmental filtering, and spatial structure on metacommunity dynamics.

    Moritz, C., Meynard, C.N., Devictor, V., Guizien, K., Labrune, C., Guarini, J.M., & Mouquet N. (2013). Disentangling the role of connectivity, environmental filtering, and spatial structure on metacommunity dynamics. Oikos, 122, 1401-1410, doi: 10.1111/j.1600-0706.2013.00377.x

  72. [64]
    Disentangling the drivers of metacommunity structure across spatial scales.

    Meynard C.M., Lavergne S., Boulangeat I., Garraud L., Van Es J., Mouquet N. and Thuiller W. (2013). Disentangling the drivers of metacommunity structure across spatial scales. Journal of Biogeography, 40, 1560-1571, doi:10.1111/jbi.12116

  73. [63]
  74. [62]
    A road map for integrating eco-evolutionary processes into biodiversity models

    Thuiller, W., Munkemuller, T, Lavergne, S., Mouillot, D., Mouquet, N., Schiffers, K. and Gravel, D. (2013).A road map for integrating eco-evolutionary processes into biodiversity models. Ecology Letters, 16, 94-105, doi:10.1111/ele.12104

  75. [61]
    Rare Species Support Vulnerable Functions in High Diversity Ecosystems.

    Mouillot, D., Bellwood, D.R., Baraloto, C., Chave, J., Galzin, R., Harmelin-Vivien, M., Kulbicki, M., Lavergne, L., Lavorel, S., Mouquet, N., Paine, C.E.T., Renaud, J. and Thuiller, T. (2013). Rare Species Support Vulnerable Functions in High Diversity Ecosystems.. PLoS Biology, 11, e1001569, doi:10.1371/journal.pbio.1001569

  76. [60]
    Unifying sources and sinks in ecology and Earth sciences

    Loreau, M., Daufresne, T., Gonzalez, A., Gravel, D., Guichard, F., Leroux, S.J., Loeuille, N., Massol, F. and Mouquet, N. (2013). Unifying sources and sinks in ecology and Earth sciences. Biological Review, 88, 365-79, doi:10.1111/brv.12003

  77. [59]
    Ecological strategies shape the insurance potential of biodiversity

    Matias, M., Combe, M., Barbera, C. and Mouquet, N. (2013). Ecological strategies shape the insurance potential of biodiversity. Frontiers in Microbiology 3: 1-9, doi:10.3389/fmicb.2012.00432

  78. [58]
    Extending the concept of keystone species to communities and ecosystems

    Mouquet N., Gravel D., Massol F. and Calcagno V. (2013). Extending the concept of keystone species to communities and ecosystems. Ecology Letters, 16, 1-8, doi:10.1111/ele.12014

  79. [57]
    Dispersal stochasticity mediates species richness in source sink metacommunities

    Matias M.G., Mouquet N. and Chase J.M. (2012). Dispersal stochasticity mediates species richness in source sink metacommunities. Oikos, 122, 395-402, doi:10.1111/j.1600-0706.2012.20479.x

  80. [56]
    Competition colonization dynamics in experimental bacterial metacommunities

    Livingston G.,Matias W.M., Calcagno C., Barbera C., Combe M., Leibold M. and Mouquet N. (2012). Competition colonization dynamics in experimental bacterial metacommunities. Nature Communication, 3, 1234, doi:10.1038/ncomms2239

  81. [55]
    The dissimilarity of species interaction networks

    Poisot T., Canard E., Mouillot D., Mouquet N. and Gravel D. (2012). The dissimilarity of species interaction networks. Ecology Letters, 15, 1353-1361, doi:10.1111/ele.12002, suppl. mat., R package

  82. [54]
    Phylogenetic constraints on ecosystem functioning

    Gravel D.*, Thomas Bell T., Claire Barbera C., Combe. M, Pommier T., and Mouquet N.* (2012). Phylogenetic constraints on ecosystem functioning. Nature Communication, 3, 1117, doi:10.1038/ncomms2123 (* These authors contributed equally to this work)

  83. [53]
    Ecophylogenetics: advances and perspectives

    Mouquet N., Devictor V., Meynard C.N., Munoz F., Bersier L.F., Chave J., Couteron P., Dalecky A., Fontaine C., Gravel D., Hardy O.J., Jabot F., Lavergne S., Leibold M., Mouillot D., Munkemuller T., Pavoine S., Prinzing A., Rodrigues A.S.L., Rohr R.P., Thebault E. and Thuiller W. (2012). Ecophylogenetics: advances and perspectives. Biological Reviews, 87, 769-785, doi:10.1111/j.1469-185X.2012.00224.x

  84. [52]
    Emergence of Structural Patterns in Neutral Trophic Networks

    Canard E., Mouquet N., Marescot L., Gaston K.J., Gravel D. and Mouillot D. (2012). Emergence of Structural Patterns in Neutral Trophic Networks. PLoS ONE, 7, e38295, doi:10.1371/journal.pone.0038295

  85. [51]
    A Phylogenetic Perspective on the Evolution of Mediterranean Teleost Fishes

    Meynard C.N., Mouillot D., Mouquet N. and Douzery E.J.P. (2012). A Phylogenetic Perspective on the Evolution of Mediterranean Teleost Fishes. PLoS ONE, 7, e36443, doi:10.1371/journal.pone.0036443

  86. [50]
  87. [49]
    A comparative study of ecological specialization estimators

    Poisot T., Canard E., Mouquet N. and Hochberg M.E. (2012). A comparative study of ecological specialization estimators. Methods in Ecology and Evolution, 3, 537-544, doi:10.1111/j.2041-210X.2011.00174.x

  88. [48]
    Using phylogenies in conservation: new perspectives

    Rolland J., Cadotte M.W., Davies J., Devictor V., Lavergne S., Mouquet N. , Pavoine S., Ana Rodrigues, Thuiller W., Turcati L., Winter M., Zupan L., Jabot F. and Morlon H. (2012). Using phylogenies in conservation: new perspectives Biology Letters, 8, 692-694, doi:10.1098/rsbl.2011.1024

  89. [47]
    From diversity indices to community assembly processes: A test with simulated data

    Munkemuller T., de Bello F., Meynard C.N., Gravel D., Lavergne S., Mouillot D., Mouquet N. and Thuiller W. (2011). From diversity indices to community assembly processes: A test with simulated data. Ecography, 34, 001-013, doi:10.1111/j.1600-0587.2011.07259.x

  90. [46]
  91. [45]
    Trophic theory of island biogeography

    Gravel D., Massol F., Canard E., Mouillot D. and Mouquet N. (2011). Trophic theory of island biogeography. Ecology Letters, 14, 1010-1016, doi:10.1111/j.1461-0248.2011.01667.x

  92. [44]
    Constraints on food chain length arising from regional metacommunity dynamics

    Calcagno V., Massol F., Mouquet N., Jarne P. and David P. (2011). Constraints on food chain length arising from regional metacommunity dynamics. Proceedings of the Royal Society of London Serie B, 278, 3042-3049, doi:10.1098/rspb.2011.0112

  93. [43]
    Empirical approaches to metacommunities: a review and comparison with theory

    Logue J.B., Mouquet N., Hannes Peter H., Hillebrand H., and The Metacommunity Working Group*. (2011). Empirical approaches to metacommunities: a review and comparison with theory. Trends in Ecology and Evolution, 26, 482-491, doi:10.1016/j.tree.2011.04.009 * Priscilla Declerck, Andreas Flohre, Stephan Gantner, Nils Gulzow, Paul Hortnagl, Sandra Meier and Bert Pecceu.

  94. [42]
    Protected and Threatened Components of Fish Biodiversity in the Mediterranean Sea

    Mouillot D., Albouy C., Guilhaumon F., Lasram F.B.R., Coll M., Devictor V., Meynard C.N., Pauly D., Tomasini J.A., Troussellier M., Velez L., Watson R., Douzery E.J.P. and Mouquet N. (2011). Protected and Threatened Components of Fish Biodiversity in the Mediterranean Sea. Current Biology, 21,1044-1050, doi:10.1016/j.cub.2011.05.005

  95. [41]
    Persistence Increases with Diversity and Connectance in Trophic Metacommunities

    Gravel D., Canard E., Guichard F. and Mouquet N. (2011). Persistence Increases with Diversity and Connectance in Trophic Metacommunities. PLoS ONE, 6(5): e19374, doi:10.1371/journal.pone.0019374

  96. [40]
    Extinction Debt in Source-Sink Metacommunities

    Mouquet N., Matthiessen B, Miller T, Gonzalez A (2011). Extinction Debt in Source-Sink Metacommunities. PLoS ONE 6(3): e17567, doi:10.1371/journal.pone.0017567

  97. [39]
  98. [38]
    Linking community and ecosystem dynamics through spatial ecology

    Massol F., Gravel D., Mouquet N., Cadotte M.W., Fukami T. and Leibold M.A. (2011). Linking community and ecosystem dynamics through spatial ecology. Ecology Letters, 14, 313-323, doi:10.1111/j.1461-0248.2011.01588.x

  99. [37]
    Experimental niche evolution alters the strength of the diversity-productivity relationship

    Gravel D., Bell T., Barbera C., Bouvier T., Pommier T., Venail P. and Mouquet N. (2011). Experimental niche evolution alters the strength of the diversity-productivity relationship. Nature, 469,89-92, doi:10.1038/nature09592

  100. [36]
    Biodiversity and Climate Change: Integrating Evolutionary and Ecological Responses of Species and Communities

    Lavergne S., Mouquet N., Thuiller W., and Ronce O. (2010). Biodiversity and Climate Change: Integrating Evolutionary and Ecological Responses of Species and Communities. The Annual Review of Ecology, Evolution, and Systematics , 41,321-350, doi:10.1146/annurev-ecolsys-102209-144628

  101. [35]
    Patch Dynamics, Persistence, and Species Coexistence in Metaecosystems

    Gravel D., Mouquet N., Loreau M., and Guichard F. (2010). Patch Dynamics, Persistence, and Species Coexistence in Metaecosystems. The American Naturalist, 176, 289-302, doi:10.1086/655426

  102. [34]
    Source and sink dynamics in metaecosystems

    Gravel D., Guichard F., Loreau M. and Mouquet N. (2010). Source and sink dynamics in metaecosystems. Ecology, 91, 2172-2184, doi:10.1890/09-0843.1

  103. [33]
  104. [32]
  105. [31]
    Defining and measuring ecological specialization

    Devictor V., Clavel J., Julliard R., Lavergne S., Mouillot D., Thuiller W., Venail P., Villeger S. and Mouquet N. (2010). Defining and measuring ecological specialization. Journal of Applied Ecology, 47, 14-25, doi:10.1111/j.1365-2664.2009.01744.x

  106. [30]
    Biodiversity as spatial insurance: the effects of habitat fragmentation and dispersal on ecosystem functioning

    Gonzalez A., Mouquet N., and Loreau M. (2009). Biodiversity as spatial insurance: the effects of habitat fragmentation and dispersal on ecosystem functioning. In: Biodiversity, Ecosystem Functioning and Ecosystem Services (eds. Naeem S, Bunker D, Hector A, Loreau M & Perrings C). Oxford University Press, Oxford

  107. [29]
  108. [28]
    Functional Diversity and Productivity Peak at Intermediate Levels of Dispersal in Evolving Metacommunities

    Venail P.A.*, MacLean R.C.*, Bouvier T., Brockhurst M.A., Hochberg M.E. and Mouquet N.* (2008) Functional Diversity and Productivity Peak at Intermediate Levels of Dispersal in Evolving Metacommunities. Nature, 452, 210-215, doi:10.1038/nature06554 (* These authors contributed equally to this study).

  109. [27]
    Evaluating Support for the Resource-ratio Hypothesis: A response to Wilson et al.

    Miller T.E., Burns J.H., Munguia P., Walters E.L., Kneitel J.M., Richards P.M., Mouquet N. and Buckley H.L. (2007). Evaluating Support for the Resource-ratio Hypothesis: A response to Wilson et al. American Naturalist.5-707-708, doi:10.1086/513110

  110. [26]
  111. [25]
    Nutrient limitation in Sarracenia purpurea microcosms

    Gray S.M., Miller T.E., Mouquet N. and Daufresne T. (2006). Nutrient limitation in Sarracenia purpurea microcosms. Hydrobiologia, 573, 173-181, doi:10.1007/s10750-006-0265-2

  112. [24]
    Which immigration policy for optimal coexistence?

    Calcagno V., Mouquet N., Jarne P., and David P. (2006). Which immigration policy for optimal coexistence?. Ecology Letters, 9, 909-911, doi:10.1111/j.1461-0248.2006.00947.x

  113. [23]
    Coexistence in a metacommunity: the competition-colonization trade-off is NOT dead

    Calcagno V., Mouquet N., Jarne P., and David P. (2006). Coexistence in a metacommunity: the competition-colonization trade-off is NOT dead. Ecology Letters, 9, 897–907, doi:10.1111/j.1461-0248.2006.00930.x

  114. [22]
  115. [21]
    Maculinea alcon

    Mouquet N., Belrose V., Thomas J.A., Elmes G.W., Clarke R.T., and Hochberg M.E. (2005). Conserving community modules: a case study of the endangered lycaenid butterfly Maculinea alcon. Ecology, 86, 3160-3173, doi:10.1890/04-1664

  116. [20]
    Maculinea arion

    Mouquet N., Thomas J.A., Elmes G.W., Clarke R.T., and Hochberg M.E. (2005). Population dynamics and conservation of a specialized predator: a case study of Maculinea arion. Ecological Monograph, 75, 525-542, doi:10.1890/05-0319

  117. [19]
    The world is patchy and heterogeneous! Trade-off and source sink dynamics in competitive metacommunities

    Mouquet N., Hoopes M.F., and Amarasekare P. (2005). The world is patchy and heterogeneous! Trade-off and source sink dynamics in competitive metacommunities. In : Metacommunities: Spatial dynamics and ecological communities. Eds M. Holyoak, M A. Leibold and R. Holt. Chicago University Press.

  118. [18]
    Metacommunities: A framework for large-scale community ecology

    Holyoak M., Leibold M.A., Mouquet N., and Holt R.D. (2005). Metacommunities: A framework for large-scale community ecology. In : Metacommunities: Spatial dynamics and ecological communities. Eds M. Holyoak, M A. Leibold and R. D. Holt. Chicago University Press.

  119. [17]
    From metacommunities to meta-ecosystems

    Loreau M., Mouquet N., and Holt R.D. (2005). From metacommunities to meta-ecosystems. In : Metacommunities: Spatial dynamics and ecological communities. Eds M. Holyoak, M A. Leibold and R. D. Holt. Chicago University Press.

  120. [16]
    Competing theories for competitive metacommunities

    Chase J., Amarasekare P., Cottenie K., Gonzalez A., Holt R., Holyoak M., Hoopes M.F, Law R., Leibold M.A., Loreau M., Mouquet N., Shurin J., and Tilman D. (2005). Competing theories for competitive metacommunities. In : Metacommunities: Spatial dynamics and ecological communities. Eds M. Holyoak, M A. Leibold and R. D. Holt. Chicago University Press.

  121. [15]
    Modelling the local population dynamics of Maculinea and their spatial interactions with their larval foodplant and Myrmica ant species

    Clarke R.T., Mouquet N., Thomas J.A., Hochberg M.E., Elmes G.W., Tesar D., Singer A. and Hale J. (2005). Modelling the local population dynamics of Maculinea and their spatial interactions with their larval foodplant and Myrmica ant species. In: Studies on the Ecology and Conservation of Butterflies in Europe. Vol. 2: Species Ecology along a European Gradient: Maculinea Butterflies as a Model, J. Settele, E. Kuhn and J.A. Thomas (Eds) pp 115-119. Pensoft, Sofia.

  122. [14]
    A Critical Review of Twenty Years' Use of the Resource-ratio Theory

    Miller T.E., Burns J.H., Munguia P., Walters E.L., Kneitel J.M., Richards P.M., Mouquet N. and Buckley H.L. (2005). A Critical Review of Twenty Years' Use of the Resource-ratio Theory. American Naturalist, 165, 439-448, doi:10.1086/428681

  123. [13]
  124. [12]
    Mechanisms of coexistence in competitive metacommunities

    Amarasekare P., Hoopes M.F., Mouquet N., and Holyoak M. (2004). Mechanisms of coexistence in competitive metacommunities. American Naturalist, 164, 310-326, doi:10.1086/422858

  125. [11]
    The metacommunity concept: A framework for multi-scale community ecology

    Leibold M.A., Holyoak M., Mouquet N., Amarasekare P., Chase J., Hoopes M., Holt R., Shurin J.B., Law R., Tilman D., Loreau M., and Gonzalez A. (2004). The metacommunity concept: A framework for multi-scale community ecology. Ecology letters, 7, 601-613, doi:10.1111/j.1461-0248.2004.00608.x

  126. [10]
    Community patterns in source-sink metacommunities

    Mouquet N. and Loreau, M. (2003). Community patterns in source-sink metacommunities. American Naturalist, 162, 544-557, doi:10.1086/378857

  127. [9]
    Community assembly time and the relationship between local and regional species richness

    [09] Mouquet N., Munguia P., Kneitel J.M. and Miller T.E. (2003). Community assembly time and the relationship between local and regional species richness. Oikos, 103, 618-626, doi:10.1034/j.1600-0706.2003.12772.x

  128. [8]
    Biodiversity as spatial insurance in heterogeneous landscapes

    [08] Loreau M., Mouquet N. and Gonzalez A. (2003). Biodiversity as spatial insurance in heterogeneous landscapes. PNAS, 100, 12765-12770, doi:10.1073/pnas.2235465100

  129. [7]
    Meta-ecosystems: a framework for a spatial ecosystem ecology

    [07] Loreau M., Mouquet N. and Holt, R. (2003). Meta-ecosystems: a framework for a spatial ecosystem ecology. Ecology Letters, 6, 673-679, doi:10.1046/j.1461-0248.2003.00483.x

  130. [6]
    Coexistence in a metacommunity: The regional similarity hypothesis

    [06] Mouquet N. and Loreau M. (2002) Coexistence in a metacommunity: The regional similarity hypothesis. American Naturalist, 159, 420-426, doi:10.1086/338996

  131. [5]
  132. [4]
  133. [3]
    The properties of competitive communities with coupled local and regional dynamics

    [03] Mouquet N., Mulder, G.S.E.E., Jansen V.A.A. and Loreau M. (2001). The properties of competitive communities with coupled local and regional dynamics. In : Dispersal: individual, population, community (Clobert, J. Danchin, E., Dhondt, A., and Nichols, J.D. Eds.): Oxford University Press, Oxford.

  134. [2]
    Coexistence, saturation and invasion resistance in simulated plant assemblages

    [02] Moore, J.L., Mouquet, N, Lawton J.H. and Loreau, M. (2001). Coexistence, saturation and invasion resistance in simulated plant assemblages. Oikos, 94, 303-314, doi:10.1034/j.1600-0706.2001.940211.x

  135. [1]
    Immigration and the maintenance of local species diversity

    [01] Loreau M. and Mouquet N. (1999). Immigration and the maintenance of local species diversity. American Naturalist 154, pp. 427-440, doi:10.1086/303252